shleshmataka :
Morphology:
Root - 5-12 cm long, 1-7 mm thick, oblique and gradually tapering, bearing a few
rootlets; external surface, dirty brown; fracture, short.
Stem - Glabrous, cylindrical, hairy, slightly branched; 10-17 cm long, 1-8 mm thick,
grooved and ribbed; basal region of branches greenish-brown, apical region dark green,
bearing a number of flowers; fracture, short.
Leaf - Simple, dark-green, smooth, alternate, opposite, exstipulate, 2.5-5 cm long,
1.8-3.6 cm broad, elliptical, lanceolate, obtuse or acutely toothed; shape and size
variable; petiole short; odour, slightly characteristic.
Histology:
Root - Mature root shows 4-5 layered cork, consisting of tabular, tangentially elongated,
thick-walled cells filled with reddish-brown contents; secondary cortex consists of a
wide zone of thin-walled, parenchymatous cells having a few resin ducts; secondary
phloem, a narrow zone, composed of sieve elements and phloem parenchyma, traversed
by phloem rays; xylem well-developed, composed of vessels, tracheids, fibres and xylem parenchyma, traversed by 1-5 seriate xylem rays; xylem vessels usually solitary
or 2-4 in groups with reticulate thickening; fibres aseptate and pointed.
Stem - Mature stem shows several bulges at places and consists of a single layered
epidermis, externally covered with a striated cuticle; a number of epidermal cells
elongate to form multicellular covering and T-shaped trichomes with 2-6 celled stalk;
cortex 3-5 layers of thin-walled, tangentially elongated parenchymatous cells, a few
filled with reddishbrown content, bulges show a few layers of collenchyma between
epidermis and parenzhymatous cortex; endodermis single layered, composed of barrelshaped cells; pericycle occurs in the form of groups of pericyclic fibres; phloem consists
of strands of sieve tubes, companion cells and phloem parenchyma; xylem consists of
vessel, parenchyma and fibres; xylem vessls show reticulate thickening; parenchyma in
abundance and paratracheal; fibres thick-walled, aseptate, short, with pointed ends;
medullary rays 2-11 cells wide; central portion occupied by pith composed of hexagonal
to polygonal, thin-walled parenchymatous cells; a few simple starch grains present in
cortical cells; cluster crystals of calcium oxalate occasionally found in pith.
Leaf -
Petiole - shows a somewhat circular outline with two lateral projections one on each
side; epidermis on both surfaces, covered externally with striated cuticle and have both
type of trichomes as described in case of stem, followed by 2-3 layers of collenchyma on
upper and lower side; stele composed of three collateral vascular bundles located in
centre, central one larger and lateral two smaller; ground tissue composed of thin-walled
parenchymatous cells, a few having oil globules and rosette crystals of calcium oxalate.
Midrib - shows similar structure as described in petiole except for 1 or 2 layers of
collenchymatous cells below both epidermis and a single vascular bundle in centre; oil
globules and rosette crystals of calcium oxalate present in a few cells of ground tissue.
Lamina - shows dorsiventral structure; epidermis single layered on either surface,
composed of thin-walled, tangentially elongated cells, covered externally with striated
cuticle; trichomes similar to those of stem; palisade single layered; spongy parenchyma
4-5 layered, loosely arranged cells; vascular bundles embedded in spongy parenchyma;
rosette crystals of calcium oxalate and oil globules present in this region; anomocytic
stomata present on both surfaces.
Dr Mohamed Rifas
A dual country licensed Ayurveda Physician, currently working in Dubai as Head of Ayurveda and Yoga department. He also works as visiting doctor for Dr Hassan Ayurveda Brain and Spine Specialty Hospital. He is specialized in joint issues, brain and spine related issues and gastrointestinal issues with respect to anxiety.
Doctor profile: Click here